Publications

google scholar

2026

Toth LT and Courtney TA. 2026. Repeated coral bleaching events have eroded reef accretion potential in the Florida Keys. Frontiers in Marine Science 13:1878462. https://doi.org/10.3389/fmars.2026.1878462

Kekuewa SAH, Rintoul MS, Courtney TA, Pezner AK, Takeda T, Kayanne H, Mitarai S, Andersson AJ. 2026. Seasonal Reef-Scale Variability in Seawater CO₂ Chemistry in the Presence and Absence of Seaweed Cultivation in Onna, Okinawa, Japan. Estuaries and Coasts 49(168). https://doi.org/10.1007/s12237-026-01770-x

Cordero E, Sherman CE, Yadav P, Kumar Raju V, Townsend JE, Courtney TA. 2026. Coral Intel: a YOLOv8 deep learning framework for monitoring Caribbean corals. Intelligent Marine Technology and Systems 4(18). https://doi.org/10.1007/s44295-026-00107-5

Infante R, Centeno L, Courtney TA, Cruz Motta JJ, Armstrong RA. 2026. Quantifying Marine Surface Microplastics in La Parguera Natural Reserve, Puerto Rico. Microplastics 5(108). https://doi.org/10.3390/microplastics5020108

DeCarlo TM, Jasnos O, Strange A, Andersson A, Bautista VII AT, Bloomer S, Bolden IW, Bosman M, Brachert TC, Braz GB, Cardoso GO, Carricart-Ganivet JP, Carilli JE, Castillo KD, Cavole L, Chan S, Chen X, Chomitz B, Correge T, Courtney TA, Deigan M, D’Olivo JP, Dunbar R, Enochs IC, Falsarella L, Felis T, Gutierrez-Estrada G, Hedger B, Hu S, Jameson S, Jupiter S, Kench P, Kersting DK, Lin K, Liu Y-W, Lorigados CAB, Manzello DP, McCulloch MT, Mies M, Moura RL, Oberle F, Pereira N, Prouty N, Ramos RD, Ren H, Ryan E, Thompson DM, Toth LT, Vergotti MJ, Webster JM, Zinke J. 2026. CoralCache: a virtual coral core repository for transparent and reproducible annual growth rate analyses, Earth System Science Data 18, 3341–3353. https://doi.org/10.5194/essd-18-3341-2026

Kekuewa SAH, Pezner AK, Courtney TA, Wei Y, Rintoul MS, Chou W-C, Soong K, Andersson AJ. 2026. Present and future seawater CO2 chemistry across multiple coral reef habitats and scales at Dongsha Atoll and Taiping Island in the South China Sea. Coral Reefs. https://doi.org/10.1007/s00338-026-02868-1

Toth LT, Borer ET, Burkepile DE, Dudney J, Lemoine NP, Renzi JJ, Smith KEL, Courtney TA, Goeking SA, Hammond WM, Hoover DL, MacFayden S, Osland MJ, Townsend JE, Fidler RY. 2026. Scenarios and strategies for future-proofing ecosystem management under climatic novelty. Conservation Biology. e70250. https://doi.org/10.1111/cobi.70250

Cornwall CE, Timmerman O, Andersson A, Berbece DM, Branson O, Comeau S, Courtney TA, Donner SD, Enochs I, Harvey BP, Isaak AL, Koch HE, Logan CA, Perry CT, Schoepf V, Toth LT, Webb A. 2026. Persistence of coral reef structures into the twenty-first century. Nature Reviews Earth & Environment. 2026 Feb 19:1-1. https://doi.org/10.1038/s43017-026-00764-4

Courtney TA, Ricaurte-Chica M, Bayo Torres AD, Chabrier-Alpi AM, Medina Martinez AG, Melendez Declet CV, Mejías-Rivera CL, Fontanez Diaz CF, Hernández Rodríguez C, Lebron-Maldanado CI, Diaz Jimenez E, Irizarry Torres I, Carballeira Martínez J, Brito-Vera MJ, Luquis Ramos I, Santos Echevarría I, Collazo Reyes JJ, Centeno Mejias LJ, Nieves-Ortíz MA, Olmeda Saldaña MF, Gordon M, Mendez Falcon V, Reyes Claudio VR, Garcia Fernandez A, Melendez Aguilar AR, Garcia Hernandez WE, Feliberty Ruiz A, Cruz-Motta JJ. Puerto Rico Coral Reef Monitoring Program Water Quality Data from 2023–2025. 2026. Scientific Data 13(150). https://doi.org/10.1038/s41597-025-06468-6

Luquis Ramos IM, Williams SM, Cruz-Motta JJ, Courtney TA. 2026. Modeling bioerosion rates by Diadema antillarum populations to improve coral reef restoration in the Caribbean. Restoration Ecology. https://doi.org/10.1111/rec.70302

Rodríguez Hernández C, Reyes JC, Perez DI, Gonzalez-Corredor J, Krause JR, Campbell JE, Fourqurean JW, Cruz Motta JJ, Courtney TA. 2026. Influence of Environmental Factors on pH Buffering, Productivity, and Blue Carbon Storage in Contrasting Thalassia testudinum Seagrass Meadows in La Parguera, Puerto Rico. Estuaries and Coasts 49(40). https://doi.org/10.1007/s12237-025-01652-8

2025

Mejias-Rivera CL, Ricaurte-Chica M, Carballeira-Martínez J, Hernández WJ, Armstrong RA, Cruz-Motta JJ, Courtney TA. 2026. Assessment of Coral Reef Eutrophication Thresholds Across Puerto Rico Using In-situ and Satellite-derived Chlorophyll-a. Estuaries and Coasts 49(26). https://doi.org/10.1007/s12237-025-01644-8

Martinez Ortiz J, Perez Perez J, Armstrong RA, Cruz Motta JJ, Courtney TA. 2025. Hypoxia driven by Caribbean Sargassum accumulation events. Royal Society Open Science 12(8):250322. https://doi.org/10.1098/rsos.250322

Courtney TA & Andersson AJ. 2025. Revisiting the terms used in net ecosystem calcification studies, carbonate budgets, and the equation of coral reef growth. Coral Reefs 44: 1031–1036. https://doi.org/10.1007/s00338-025-02643-8

Sanassy Pilly S, Townsend JE, Alisa CA, Razak TB, Roche RC, Turner JR, Chan S, Kriegman DJ, Andersson AJ, Perry CT, Lange ID, Courtney TA. 2025. Quantifying coral reef carbonate budgets: a comparison between ReefBudget and CoralNet. Coral Reefs 44: 513–527. https://doi.org/10.1007/s00338-025-02620-1

2024

Pezner AK, Courtney TA, Chou W-C, Chu H-C, Frable BW, Kekuewa SAH, Soong K, Wei Y, Andersson AJ. 2024. Coral growth along a natural gradient of seawater temperature, pH, and oxygen in a nearshore seagrass bed on Dongsha Atoll, Taiwan. PLoS ONE 19(10): e0312263. https://doi.org/10.1371/journal.pone.0312263

Mejias-Rivera CL & Courtney TA. 2024. Ocean Warming, Heat Stress, and Coral Bleaching in Puerto Rico. Caribbean Journal of Science (54): 132-149. https://doi.org/10.18475/cjos.v54i1.a17

2023

Toth LT, Courtney TA, Colella M, Ruzicka RR. 2023. Stony coral tissue loss disease accelerated shifts in coral composition and declines in reef accretion potential in the Florida Keys. Frontiers in Marine Science. https://doi.org/10.3389/fmars.2023.1276400

Mejias-Rivera CL, Armstrong RA, Balint S, García-Troche E, McKinney RA, Morell JM, Oczkowski A, Courtney TA. 2023. Localized inshore warming, acidification, and elevated particulate organic matter across a coupled mangrove, seagrass, and coral reef ecosystem in La Parguera, Puerto Rico. Coral Reefs. https://doi.org/10.1007/s00338-023-02435-y

Cornwall CE, Carlot J, Branson O, Courtney TA, Harvey BP, Perry CT, Andersson AJ, Diaz-Pulido G, Johnson MD, Kennedy E, Krieger EC, Mallela J, McCoy SJ, Nugues MM, Quinter E, Ross CL, Ryan E, Saderne V, Comeau S. 2023. Crustose coralline algae can contribute more than corals to coral reef carbonate production. Nature Communications Earth & Environment. https://doi.org/10.1038/s43247-023-00766-w

Pezner AK, Courtney TA, Barkley HC, Chou WC, Chu HC, Clements SM, Cyronak T, DeGrandpre MD, Kekuewa SAH, Kline DI, Liang Y-B, Martz TB, Mitarai S, Page HN, Rintoul MS, Smith JE, Soong K, Takeshita Y, Tresguerres M, Wei Y, Yates KK, Andersson AJ. 2023. Increasing hypoxia on global coral reefs under ocean warming. Nature Climate Change 13, 403–409 https://doi.org/10.1038/s41558-023-01619-2

2022

Rintoul M, Courtney T, Dohner J, Giddings S, Inoha K, Kekuewa S, Mitarai S, Monismith S, Pezner A, Andersson A. 2022. The Effects of Light Intensity and Flow Speed on Biogeochemical Variability within a Fringing Coral Reef in Onna-son, Okinawa, Japan. Journal of Geophysical Research – Oceans. https://doi.org/10.1029/2021JC018369

Kekuewa SAH, Courtney TA, Cyronak T, Andersson AJ. 2022. Seasonal nearshore ocean acidification and deoxygenation in the Southern California Bight. Scientific Reports. https://doi.org/10.1038/s41598-022-21831-y

Toth LT, Courtney TA, Colella MA, Kupfner Johnson SA, Ruzicka RR. 2022. The past, present, and future of coral reef growth in the Florida Keys. Global Change Biology. https://doi.org/10.1111/gcb.16295

Courtney TA, Barkley HC, Chan S, Couch CS, Kindinger TL, Oliver TA, Kriegman DJ, Andersson AJ. 2022. Rapid assessments of Pacific Ocean net coral reef carbonate budgets and net calcification following the 2014-2017 global coral bleaching event. Limnology and Oceanography. https://doi.org/10.1002/lno.12159

Elahi R, Edmunds PJ, Gates RD, Kuffner IB, Barnes BB, Chollett I, Courtney TA, Guest JR, Lenz EA, Toth LT, Viehman TS, Williams ID. 2022. Scale dependence of coral reef oases and their environmental correlates. Ecological Applications. https://doi.org/10.1002/eap.2651

Bresnahan P, Cyronak T, Brewin RJW, Andersson A, Wirth T, Martz T, Courtney T, Hui N, Kastner R, Stern A, McGrain T, Reinicke D, Richard J, Hammond K, Waters S. 2022. A High-Tech, Low-Cost, Internet of Things Surfboard Fin for Coastal Citizen Science, Outreach, and Education. Continental Shelf Research. https://doi.org/10.1016/j.csr.2022.104748

2021

Courtney TA, Cyronak T, Griffin AJ, Andersson AJ. 2021. Implications of salinity normalization of seawater total alkalinity in coral reef metabolism studies. PLOS ONE. https://doi.org/10.1371/journal.pone.0261210

Kekuewa SAH, Courtney TA, Cyronak T, Kindeberg T, Eyre BD, Stoltenberg L, Andersson AJ. 2021. Temporal and Spatial Variabilities of Chemical and Physical Parameters on the Heron Island Coral Reef Platform. Aquatic Geochemistry. https://doi.org/10.1007/s10498-021-09400-7

Courtney TA, Guest JR, Edwards AJ, Dizon RM. 2021. Linear extension, skeletal density, and calcification rates of the blue coral Heliopora coerulea. Coral Reefs. https://doi.org/10.1007/s00338-021-02137-3

Pezner AK, Courtney TA, Page HN, Giddings SN, Beatty CM, DeGrandpre MD, Andersson AJ. 2021. Lateral, Vertical, and Temporal Variability of Seawater Carbonate Chemistry at Hog Reef, Bermuda. Frontiers in Marine Science. https://doi.org/10.3389/fmars.2021.562267

2020

Courtney TA, Kindeberg T, Andersson AJ. 2020. Coral calcification responses to the North Atlantic Oscillation and coral bleaching in Bermuda. PLOS ONE. https://doi.org/10.1371/journal.pone.0241854

Kindeberg T, Bates NR, Courtney TA, Cyronak T, Griffin A, Mackenzie FT, Paulsen M-L, Andersson AJ. 2020. Porewater Carbonate Chemistry Dynamics in a Temperate and a Subtropical Seagrass System. Aquatic Geochemistry. https://doi.org/10.1007/s10498-020-09378-8

Courtney TA, Barnes BB, Chollett I, Elahi R, Gross K, Guest JR, Kuffner IB, Lenz EA, Nelson HR, Rogers CS, Toth LT, Andersson AJ. 2020. Disturbances drive changes in coral community assemblages and coral calcification capacity. Ecosphere. https://doi.org/10.1002/ecs2.3066

Cyronak T, Takeshita Y, Courtney TA, DeCarlo EH, Eyre BD, Kline DI, Martz T, Page H, Price NN, Smith J, Stoltenberg L, Tresguerres M, Andersson AJ. 2020. Diel temperature and pH variability scale with depth across diverse coral reef habitats. Limnology and Oceanography Letters. https://doi.org/10.1002/lol2.10129

2019

Baumann JH, Ries JB, Rippe JP, Courtney TA, Aichelman HE, Westfield I, Castillo KD. 2019. Nearshore coral growth declining on the Mesoamerican Barrier Reef System. Global Change Biology, 25: 3932–3945. https://doi.org/10.1111/gcb.14784

Courtney TA & Andersson AJ. 2019. Evaluating measurements of coral reef net ecosystem calcification rates. Coral Reefs, 38(5):997–1006. https://doi.org/10.1007/s00338-019-01828-2

Page HN, Courtney TA, De Carlo EH, Howins NM, Koester I, Andersson AJ. 2019. Spatiotemporal variability in seawater carbon chemistry for a coral reef flat in Kāne’ohe Bay, Hawai’i. Limnology and Oceanography, 63:913–934. https://doi.org/10.1002/lno.11084

2018

Guest JR, Edmunds PJ, Gates RD, Kuffner IB, Andersson AJ, Barnes BB, Chollett I, Courtney TA, Elahi R, Gross K, Lenz EA, Mitarai S, Mumby PJ, Nelson HR, Parker BA, Putnam HM, Rogers CS, Toth LT. 2018. A framework for identifying and characterising coral reef “oases” against a backdrop of degradation. Journal of Applied Ecology, 00:1-11. https://doi.org/10.1111/1365-2664.13179

Courtney TA, De Carlo EH, Page HN, Bahr KD, Barro A, Howins N, Tabata R, Terlouw G, Rodgers KS, Andersson AJ. 2018. Recovery of reef-scale calcification following a bleaching event in Kāne’ohe Bay, Hawai’i. Limnology & Oceanography Letters, 3:1–9. https://doi.org/10.1002/lol2.10056

2017

Courtney TA, Lebrato M, Bates NR, Collins A, de Putron SJ, Garley R, Johnson, R, Molinero JC, Noyes TJ, Sabine CL, Andersson AJ. 2017. Environmental controls on modern scleractinian coral and reef-scale calcification. Science Advances, 3(11), p.e1701356. https://doi.org/10.1126/sciadv.1701356

Page HN, Courtney TA, Collins A, De Carlo EH, Andersson AJ. 2017. Net community metabolism and seawater carbonate chemistry scale non-intuitively with coral cover. Frontiers in Marine Science, 4:161. https://doi.org/10.3389/fmars.2017.00161

2016

Courtney TA, Andersson AJ, Bates NB, Collins A, Cyronak T, de Putron SJ, Eyre BD, Garley R, Hochberg EJ, Johnson R, Musielewicz S, Noyes T, Sabine CL, Sutton AJ, Toncin J, Tribollet A. 2016. Comparing Chemistry and Census-based Estimates of Net Ecosystem Calcification on a Rim Reef in Bermuda. Frontiers in Marine Science, 3:181. https://doi.org/10.3389/fmars.2016.00181

Baumann JH, Townsend JE, Courtney TA, Aichelman HE, Davies SW, Lima FP, Castillo KD. 2016. Temperature Regimes Impact Coral Assemblages along Environmental Gradients on Lagoonal Reefs in Belize. PLoS ONE, 11(9): e0162098. https://doi.org/10.1371/journal.pone.0162098

Aichelman HE, Townsend JE, Courtney TA, Baumann JH, Davies SW, Castillo KD. 2016. Heterotrophy mitigates the response of the temperate coral Oculina arbuscula to temperature stress. Ecology and Evolution, 6(18): 6758-6769. https://doi.org/10.1002/ece3.2399

Horvath KM, Castillo KD, Armstrong P, Westfield IT, Courtney T, Ries JB. 2016. Next-century ocean acidification and warming both reduce calcification rate, but only acidification alters skeletal morphology of reef-building coral Siderastrea siderea. Scientific Reports, 6:29613. https://doi.org/10.1038/srep29613

2015

Courtney T and Ries JB. 2015. Impact of atmospheric pCO2, seawater temperature, and calcification rate on the δ18O and δ13C composition of echinoid calcite (Echinometra viridis). Chemical Geology, 411: 228-239. https://doi.org/10.1016/j.chemgeo.2015.06.030

2013

Courtney T, Ries JB, Westfield I. 2013. Predicted end of 21st century CO2-induced ocean acidification decreases calcification rates in the tropical urchin Echinometra viridis. Journal of Experimental Marine Biology and Ecology, 440: 169-175. https://doi.org/10.1016/j.jembe.2012.11.013